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Question 114·200 Super-Hard SAT Reading Questions·Expression of Ideas

Researchers investigating drought tolerance in wild sunflowers found that plants carrying a particular DNA variant near a water-transport gene survived dry conditions at higher rates. Because nearby variants are often inherited together, however, this field association could not identify the particular variant responsible. The researchers later altered only the candidate variant in otherwise genetically identical plants and observed increased drought survival. This experiment therefore strengthens the case that the candidate variant itself contributes to drought tolerance. ______ it shows that changing the variant can affect survival when genetic background is held constant; it does not establish how strongly the variant affects survival amid the environmental variation of natural populations.

Which choice completes the text with the most logical transition?