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Question 177·Hard·Form, Structure, and Sense

Fossil feathers rarely retain their original pigments, but microscopic structures can preserve clues to color. The recovery of tiny mineral grains from the feathers of several fossil birds, together with new analyses showing how those grains scatter light, ______ enabled researchers to model hues that would otherwise be lost to time.

Which choice completes the text so that it conforms to the conventions of Standard English?