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Question 146·Medium·Form, Structure, and Sense

Some desert plants can draw moisture from fog. In a recent laboratory study, a group of botanists _____ to determine whether tiny grooves on cactus spines guide water toward the plant’s stem placed artificial spines with different surface patterns inside a fog chamber. Grooved spines delivered more water to their bases than smooth spines did.

Which choice completes the text so that it conforms to the conventions of Standard English?