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Question 80·Medium·Boundaries

In her latest article, historian Laila Garner argues that the "Pine Needle Letters"—some of the oldest pieces of correspondence ever discovered—were preserved by an accident of climate. The letters’ ______ thin, fibrous layers of processed cedar bark—remained dry for centuries in the arid caves of the Anatolian plateau.

Which choice completes the text so that it conforms to the conventions of Standard English?