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Question 112·200 Super-Hard SAT Reading Questions·Standard English Conventions

Mycologist Nan Shi meticulously catalogs each fungal specimen she encounters on her month-long expeditions, noting spore size, metabolic by-products, and DNA _____ painstaking recordkeeping, she argues, will allow future researchers to trace how climate change alters underground networks.

Which choice completes the text so that it conforms to the conventions of Standard English?