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Question 102·Hard·Inferences

In a 2023 study, ornithologist Lydia Klimczak and colleagues examined satellite-tracking data from 148 sand martin (Riparia riparia) individuals collected between 1980 and 2020. They found that

  • the birds’ departure dates from their sub-Saharan wintering grounds remained essentially constant over the 40-year interval,
  • average air temperatures at those wintering grounds also showed no consistent trend,
  • mean spring temperatures at several key stopover wetlands along the birds’ migration corridor through North Africa and southern Europe rose by roughly 1.3C1.3\,^{\circ}\mathrm{C}, and
  • the total time spent resting at those stopovers declined by about 30 percent. As a result, sand martins reached their European breeding sites roughly seven days earlier in 2020 than in 1980. From these findings, Klimczak and colleagues inferred that over the study period, ______.

Which choice most logically completes the text?